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What is the ` (the character under the tilde) called, and how can it be used in PHP programing?

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That is a backtick (or backquote). Anything inside the backquote will be "executed" and the results may be stored in a variable. Consider this piece of code: <?php$output = `ls -al`;echo "<pre>$output</pre>";?> the ls -al command will be executed by the shell and the results are...
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That is a backtick (or backquote).  

Anything inside the backquote will be "executed"  and the results may be stored in a variable. 

Consider this piece of code:

<?php
$output = `ls -al`;
echo "<pre>$output</pre>";
?>

 

the ls -al command will be executed by the shell and the results are stored in the $output variable

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The character is called a backtick. I’m not familiar with PHP but the backtick is used commonly in a lot of programming languages.
Comments

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